5.1 The Simple Birth Process

Each individual in a population independently gives birth to one new individual in \((t, t + h]\) with probability \(\lambda h + o(h)\), and there are no deaths. Let \(X(t)\) be the population size, with \(X(0) = a\). With \(n\) individuals, the probability of exactly one birth in \((t, t + h]\) is \(n\lambda h + o(h)\), and of more than one is \(o(h)\). So \[P_n(t + h) = P_n(t)(1 - n\lambda h) + P_{n-1}(t)(n - 1)\lambda h + o(h),\] and letting \(h\to 0\), \begin {equation} \label {eq:birthde} P_n'(t) = -n\lambda P_n(t) + (n - 1)\lambda P_{n-1}(t),\qquad n\geq a . \end {equation}

Theorem 5.1.1 (Yule process). For the simple birth process with \(X(0) = a\), \(X(t)\) has the negative binomial distribution \[P(X(t) = n) = \binom {n - 1}{a - 1}e^{-a\lambda t}\left (1 - e^{-\lambda t}\right )^{n - a},\qquad n = a, a + 1, \dots ,\] with \(E\left [X(t)\right ] = ae^{\lambda t}\) and \(\var \left [X(t)\right ] = ae^{\lambda t}\left (e^{\lambda t} - 1\right )\).

Proof. Let \(M(\theta , t) = E\left (e^{\theta X(t)}\right ) = \sum _ne^{n\theta }P_n(t)\), so that \(M(\theta , 0) = e^{a\theta }\), \(\frac {\partial M}{\partial \theta } = \sum _nne^{n\theta }P_n(t)\) and \(\frac {\partial M}{\partial t} = \sum _ne^{n\theta }P_n'(t)\). Multiplying (5.1) by \(e^{n\theta }\) and summing over \(n\), \[\frac {\partial M}{\partial t} = -\lambda \frac {\partial M}{\partial \theta } + \lambda e^\theta \frac {\partial M}{\partial \theta }, \qquad \text {that is}\qquad \frac {\partial M}{\partial t} + \lambda (1 - e^\theta )\frac {\partial M}{\partial \theta } = 0 .\] This first-order equation says that \(M\) is constant along the characteristic curves \(\frac {dt}{1} = \frac {d\theta }{\lambda (1 - e^\theta )}\). Writing \(\lambda \,dt = \frac {d\theta }{1 - e^\theta } = \frac {e^{-\theta }\,d\theta }{e^{-\theta } - 1}\) and integrating, \(e^{\lambda t}(e^{-\theta } - 1)\) is constant along each curve, so \(M(\theta , t) = \Phi \left (e^{\lambda t}(e^{-\theta } - 1)\right )\) for some function \(\Phi \). At \(t = 0\), \(\Phi (e^{-\theta } - 1) = e^{a\theta }\); putting \(y = e^{-\theta } - 1\) gives \(\Phi (y) = (1 + y)^{-a}\). Hence \[M(\theta , t) = \left [1 - e^{\lambda t}\left (1 - e^{-\theta }\right )\right ]^{-a} .\] A negative binomial variable counting the trials needed for \(r\) successes with success probability \(p\) has m.g.f. \(\left (\frac {pe^\theta }{1 - (1 - p)e^\theta }\right )^r = \left [1 - \frac 1p(1 - e^{-\theta })\right ]^{-r}\), which is \(M(\theta , t)\) with \(r = a\) and \(p = e^{-\lambda t}\). The mean and variance are those of this distribution, \(\frac rp = ae^{\lambda t}\) and \(\frac {r(1 - p)}{p^2} = ae^{\lambda t}(e^{\lambda t} - 1)\). □

The mean grows exponentially, as in the deterministic model \(x'(t) = \lambda x(t)\); the randomness shows in the variance, which is of the same order as the square of the mean.

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